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Betta
coccina Vierke, 1979
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Betta
coccina
is a small red betta found in Jambi (Sumatra), southern Johor
and Malakka - in Malakka the species is now extinct - (Malaysia)
and was first described by Jörg Vierke in 1979. The name
Vierke gave the small Betta, coccina, means wine-red or
claret, a description that perfectly suits the fish if you look
at its color. The only difference between the species from Malaysia
and B. coccina from Sumatra are little color differences
which are common for most type localities.
Betta
coccina was the first fish of the B. coccina group
to be discovered and a true revolution since we only knew B.
splendens varieties with such a beautiful red color. Plus
this species had another unique mark which will be described later.
This
species habitats peat swamps that lie deep in the forests. These
swamps have extreme water conditions, fish have been found in
water with a pH between 4.0 - 6.0, a dH between 0.0 - 4.0 and
a KH of 0.0 with the water temperature being around 24° to
27°C. The reason why the temperatures are that low compared
to the savannah habitats of B. smaragdina and B. splendens
is because the puddles, in which Betta coccina lives are
surrounded by trees that stop almost any direct sunlight resulting
in shaded, cool water. The water is tea-colored as a result of
leaf litter that falls in the water providing the water conditions
described above. In water this soft and acidic there's not much
chance for plants to survive with the exception of the few Cryptocoryne
species.
These habitats are known for drying up during the dry season when
there is no rain. All that's left are small puddles and wet layers
of leafs both in which this species manages to survive until the
next rainfall. This is characteristic for the B. coccina
group.
Description
B.
coccina reaches a total length of 5,6 cm, both sexes share
that length although females can be a little smaller than males.
The first description refers to the male. The body is sometimes
slender and sometimes more B. smaragdina shaped, it depends
on the individual fish. The color of the body can differ from
light to dark brown but also from bright to wine-red. It's common
that male shows a dark brown body with two horizontal lighter
brown bars running from the eye to the start of the caudal fin.
Their eyes are bright blue. The dorsal, anal, caudal and pelvic
fins are bright to wine-red and in adults the dorsal and caudal
fins have a white edge which is black in the anal and pelvic fins.
The quills are gold to red. Both sexes possess wine-red, blackish
spots in the dorsal and caudal fin that sometimes also has white
spots in male specimens. The pectoral fins are transparent. The
unique mark that this species and a lot of other species of the
B. coccina group have is a bright blue blotch on their
sides! Only young males possess such a spot and as the male grows
older the blotch slowly disappears. Now and then the blotch may
also chance in a blue, bar like figure. However the spot is not
a rule for Betta coccina, there are a lot of recordings
where the males did not have the spot and it's even been described
some males have the spot on one side of their body but not on
the other! Conspicuous is the fact that the lateral blotch is
a spot where other B. coccina target their attack on, mostly
just gentle pushes with the mouth.
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B.
coccina females. Pictures by
Jirapat Chunthapong.
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Females
are duller versions of the males, shorter fins and less colorful
but by all means not less beautiful when ready to spawn. Their
horizontal stripe pattern, a female her most common pattern, runs
from the beginning of the caudal fin over the whole body and through
the eye and on the lips, creating a beautiful mask like image.
Males also show this mask like image though rarely. I don't think
it's necessary to go in deeper on the species their appearance,
the pictures speak for themselves.
This
species is not a very peaceful one but several males can be kept
together. They do tend to be even more aggressive when the male
is constructing a nest or when they are spawning and of course
after. In the two latter the female becomes more aggressive also,
sometimes even more than the male which is kind of uncommon since
it's in the male's character of most betta species.
This
species can be very shy and if so tends to hide all the time and
rarely come out but there are a few tricks to reverse that which
will be discussed later on.
Differentiation
from Betta livida (There is a correct revision
of this part in my coming Betta livida article.)
B.
coccina looks a lot like B. livida which is from Selangor
and the latter has often been mistaken for Betta coccina.
The reason why can be noted from the pictures of both species.
However there are small differences between the two species, mainly
coloration differences. In B. livida both sexes possess
the lateral blue blotch and in B. coccina only the male
has such a blotch plus in the latter named species it's easy to
tell whether it's a male or a female while in the first named
species both sexes look so alike it's close to impossible to determine
the sex based on coloration and fin length. Another, however minor,
difference is that B. coccina has black tipped pelvic fins
where B. livida has greenish, white tipped pelvic fins
and the species grows somewhat smaller, 5 cm, than B. coccina.
What also helps to differentiate Betta livida from B.
coccina and other species is that B. livida has two
vertical gold bars on the operculum. Both species seem also to
be different in their breeding behavior. B. livida is easier
to spawn and cannibalism has not been witnessed in B. livida.
Another conspicuous difference is that both parents tend to the
fry for the first few days.
The
B. coccina tank
Betta
coccina requires more attention than B. splendens.
I'm aiming towards the water conditions that must be maintained
and monitored at least twice a week to make sure the water has
the right pH, dH and KH values, and if the water is not polluted
what can result in death on longer terms and disease in shorter
terms. Water changes with prepared water, meaning that the same
conditions are gained as the water in the tank, should be carried
out at most once a week or at least once in two weeks with caution.
The amount of water to be changed should be around 20% to 30%.
All the values mentioned earlier will do. B. coccina
are not very active fish so a small sized tank will do fine.
Personally I use 60 x 30 x 30 cm tanks for all my Betta
species. Height of the tank is very important because this small
red Betta needs to come to the surface for air and it's
best to make his swim as short as possible. Remember that Betta
coccina lives in shallow puddles sometimes less than 20
cm deep! A tank with 30 cm in height is the maximum for me.
Also make sure the tank is covered because they are great jumpers!
The tank can do without a filter when you do regular water changes
but even if you do I still recommend using a small, slow running
filter. All that there needs to be in the filter is highly active
peat granulate and a layer of filter wool to catch the dirt.
The
tank itself must resemble their natural habitat, meaning to
heavily plant it and dim the light as much as possible. Instead
of heavily planting the tank but filling it with leaf litter
would be more natural but I'm not too fond of that because the
leafs will start to decompose within a week or two. That means
the leaf litter constantly has to be replaced by new litter
what will disturb the fish. Instead, a heavily planted tank
will do great as a substitute with maybe a few oak leafs where
one likes it. Betta coccina likes to stay near plant
cover and will try to avoid the open water as much as possible.
To create as much darkness as possible the best thing to use
at the bottom of the tank is black gravel and make sure the
back of the tank also has a black background, all this will
make B. coccina feel good and a little less shy. The
plants that can be used form only a small list; Cryptocoryne
species, Microsorium pteropus, Anubias species,
Ceratopteris pteridiodes, Ceratopteris thalictroides,
Vesicularia dubyana and on the surface Salvinia natas,
Salvinia auriculata and Riccia fluitans will do
great. Bog- and driftwood can also be used and gives a very
natural effect if placed effective.
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A
young B. coccina male displaying. Picture by Horst
Linke.
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If
such an environment is created the species will show their most
beautiful colors and their most interesting behavior. In hardly
decorated tanks they will appear with diminished colors and only
their fins will be colored red with the lateral blotch hardly
being perceptible. They will often remain in corners or near the
closest thing to cover. The tank will hardly be an encouragement
to spawn. But kept under the right circumstances it is one of
the most interesting species known and if all goes well, as described
in the breeding chapter, they will spawn.
Betta
coccina likes to lean on plant leafs, a piece of wood or sometimes
just holding still in open water without moving a fin in any of
the mentioned situations. They hold so absolutely still they can
appear to have died unless a closer look will be taken. You can
find B. coccina throughout the entire tank. Near the bottom,
where they like to lean on the gravel like they do on plant leafs,
but also in between floating plants near the surface. This species,
as all other betta species, form territories after a few days.
That is, the males form territories. Until that time B. coccina
could be hardly described as aggressive, they chase each other
now and then but it won't come to real damages to the scales and
fins. Only pushing each other with the mouth and occasionally
displaying can be observed. That changes as soon as the males
have formed their territories, more serious chases take place
and the occasional bites, yet nothing that will result in severe
damages or worse. A simple flare or look in the other's direction
is usually enough to scare the other away.
These
red Bettas eat black and white mosquito larvae, Artemia
and Tubifex. Preferably live foods but frozen will also
be taken without any problems. Tubifex is a food you should
be careful with to prevent the fish from choking and make them
fat, even though it's their main menu in their natural habitat.
I advice to feed them carefully in small amounts per fish and
monitor them instead of stuffing each fish full of food. That
way you; a) prevent leftovers from rotting in the tank and b)
each fish will get enough food individually. Also maintain a scheduled
day when the fish are not fed, it's good for their health and
their system. They approach their food very slow to suddenly take
a dive at it. Immediately after they captured the food they dive
back into the plant cover waiting for more.
Breeding Betta coccina
B.
coccina is a very difficult species to breed. The problem
lies in the fact that the pairs select themselves, and if they
don't like any of the other fish nothing will happen. Because
of this it's recommended to keep a few fish together until they
form pairs and then they can be separated from the others and
put in a breeding tank. But even when good pairs eventually form
the whole process may take several months!
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Adult
B. coccina male. Picture by Kei Sasaki.
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To
bring Betta coccina in spawning condition they must be
fed well with foods described above and here also, pay close attention
to the water its condition and values.
If
all is going well the pair can be placed in a breeding tank that
is at least 40 x 15 x 15 cm but larger is of course no problem.
The breeding tank doesn't need as much decoration as their normal
tanks, a few plants and some floating will do just fine. It is
advised to create some hiding places so they can hide from one
another. B. coccina is a bubblenester and the male builds
his nest under floating leafs, floating plants or in the tank
placed tubes. The little trick that was mentioned earlier is to
place one or more of these items in the front of the tank, that
way the fish will show themselves instead of hiding most of the
time if you have shy specimens.
If
the fish are in spawning condition the female will show her brightest
colors and won't differ much from the male, that is, with some
specimens. Her now red body will show four to five dark vertical
bars and a lighter horizontal bar that runs from the beginning
of the head all the way to the beginning of the caudal fin. A
small white genital papilla will also be present. This coloration
can be seen before the actual spawning. During spawning the female's
color is more red brownish without any bars on the flank but with
bright greenish scales here and there. From the top of the head
to the dorsal fin and from there to the caudal fin runs a white
bar as if there are no scales, chased B. coccina display
the same bar. The male keeps the same color whether he is constructing
the nest or to the point of actual spawning. The only difference
between spawning colors and normal colors are the bright green
bluish scales of the male, with the blue spot, if he possesses
one, being brighter than normal and than the scales at this moment.
It may also occur that the fins have a brighter red color than
usual.
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B.
coccina spawning, on the left a good image of the male.
On the right the female. Pictures by Kei Sasaki.
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Left:
Male embracing the female. Right: The female in the rigid
mating stance. Pictures by Kei Sasaki.
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Ready
to spawn females can be become aggressive towards a male that
is not ready even though they form a good pair. In an article
by Jörg Vierke it is explained that such a female could even
attack or kill the male. Before and during spawning males are
very aggressive towards the female what results in damage to the
female's scales and fins. Yet the female does not seem to mind
and doesn't leave the nest. This aggressive behavior by the male
occurs after numerous pushes on the side of female B. coccina
with the mouth to induce spawning. This particular behavior occurs
in both sexes while spawning, the female will gently push the
male's blotch with the mouth to induce spawning. If all goes well
the male will be at his nest as the female approaches him, or
if not, she lures him over to the nest. The two fish may swim
around each other for a while of spread their fins and send waves
to one another as a sign they are ready. The male embraces the
female on the traditional Betta way. This only lasts a
few seconds and the male ends the embrace while the female stays
in the rigid mating stance for a few more seconds. Meanwhile the
male looks for eggs which he will bring to the nest. The bubblenest
of B. coccina isn't very large what can be seen on the
picture beneath. This species is not very prolific and the number
of eggs in total is small, 20 to 50 eggs, as is the number that
a female releases at one time. The latter differs from one to
three eggs. The eggs themselves are white and rather large like
eggs of B. bellica, B. simorum and mouthbrooders.
This is the case in more species of the B. coccina group.
After spawning the female gets chased away by the male who will
take care of the eggs and it's wise to remove the female at this
stage. After 36 hours the eggs start to hatch. If the nest is
built on the underside of a floating leaf or underneath floating
plants it might happen the eggs won't hatch. Then it's advised
to darken the area above the nest to make them hatch the next
time. Now most species from the B. coccina group are known
for leaving the fry alone and for good parental care for their
fry in the free swimming stage. B. coccina does usually
not live up to that expectation so it's best to either remove
male also or relocate the fry to a small tank when they reach
the free swimming stage, which is reached in two to three days.
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B.
coccina bubblenest and eggs. Picture by Kei Sasaki.
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Raising
the fry
After
taking the fry to a small tank with some floating plants and Vesicularia
dubyana they'll go in search of food. The first three days
they will find enough themselves and on the fourth day freshly
hatched Artemia salina can be offered to the fry. The young
don't grow very fast and reach a length of 2,5 cm in 140 days.
Water
changes in the fry tank must be carried out very careful otherwise
this could lead to significant losses. If there are different
batches of fry of different ages in the same tank the older fry
must be separated from the younger fry when the older fry reaches
a length around 3,8 cm, because unlike others from the group this
species their fry do eat their younger brothers and sisters.
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Betta
coccina male. Picture by Arend van den Nieuwenhuizen.
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Other
species of the B. coccina group (Newly discovered
species are left out.)
All water conditions were monitored in the species their natural
habitats.
Betta
rutilans Witte & Kottelat, 1991
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Two
B. rutilans males. Pictures by Horst Linke and Jirapat
Chunthapong.
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| Locality |
Anjungan |
| Water
conditions |
pH
4.5, gH 1.0, kH 1.0, temp 22° - 26°C. |
| Maximum
size |
3,5
cm |
| Behavior |
Very
aggressive to other fish and they hate their own kind. |
| Breeding |
Difficult.
Bubblenester. |
| Picture
by |
Horst
Linke and Jirapat Chunthapong |
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Betta
brownorum Witte & Schmidt, 1992
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Left:
B. brownorum male. Picture by Dr. Jürgen Schmidt.
Right: A female. Picture by Eric Naus.
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| Locality |
Sibu,
Matang, West Kalimantan |
| Water
conditions |
pH
< 5.0, very soft water, temp 22° - 26°C. |
| Maximum
size |
6
cm |
| Behavior |
Peaceful
in general. |
| Breeding |
Bubblenester.
Mouthbrooding has also been reported. |
| Picture
by |
Dr.
Jürgen Schmidt and Eric Naus |
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Not
yet described species. Left: B. sp. cf. rutilans
"Green." Right: B. sp. aff. burdigala
"Pangkalanbun." Pictures by Kei Sasaki.
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Left
and right: another new species, B. sp. aff. burdigala
"Sukadana" from Kalimantan. Pictures by Team Borneo.
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Betta
livida Ng & Kottelat, 1992
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B.
livida male. Picture by Raffles Museum of
Biodiversity Research, Singapore.
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| Locality |
Selangor,
Perak |
| Water
conditions |
pH
3.5 - 3.7, soft water, temp 22° - 26°C. |
| Maximum
size |
5
cm |
| Behavior |
Peaceful
in general. |
| Breeding |
Bubblenester. |
| Picture
by |
Raffles
Museum of Biodiversity Research |
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Betta
tussyae Schaller, 1985
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A
male B. tussyae. Picture by Horst Linke.
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| Locality |
Kuantan,
Chukai, Rompin |
| Water
conditions |
pH
4.0 - 5.0, gH 1.0 - 2.0, temp 21° - 24°C. |
| Maximum
size |
5,5
cm |
| Behavior |
Peaceful
in general. |
| Breeding |
Bubblenester. |
| Picture
by |
Horst
Linke |
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Betta
burdigala Kottelat & Ng, 1994
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Male
B. burdigala. Picture by Michael Schlüter.
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| Locality |
Banka,
Kubu |
| Water
conditions |
pH
4.5, gH 0.0 - 1.0, kH 0.0 - 1.0, temp 27°C. |
| Maximum
size |
5
cm |
| Behavior |
Peaceful
in general. |
| Breeding |
Bubblenester. |
| Picture
by |
Michael
Schlüter |
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Betta
miniopinna Tan & Tan, 1994
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Left
and right: B. miniopinna males. Pictures by Kei Sasaki.
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| Locality |
Banka |
| Water
conditions |
pH
5.8 - 6.5, soft water. |
| Maximum
size |
3,2
cm |
| Behavior |
Aggressive. |
| Breeding |
Difficult.
Bubblenester. |
| Picture
by |
Kei
Sasaki (Betta House) |
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Betta
persephone Schaller, 1987
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B.
persephone. Picture by Arend van
den Nieuwenhuizen.
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| Locality |
Ayer
Hitam, Muar |
| Water
conditions |
pH
7.0, dH 10.0, temp 23° - 28°C. |
| Maximum
size |
3,2
cm |
| Behavior |
Aggressive. |
| Breeding |
Bubblenester. |
| Picture
by |
Arend
van den Nieuwenhuizen |
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Betta
coccina male. Picture by Jirapat Chunthapong.
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Pictures
by
(In order of display)
(The pictures may not be reproduced in any
way without the photographer's permission.)
Picture 1: B. coccina male, taken by Horst Linke
Picture 2: B. coccina females, taken by Jirapat Chunthapong
Picture 3: B. coccina young male, taken by Horst Linke
Picture 4: B. coccina male, taken by Kei Sasaki (Betta
House)
Picture 5: B. coccina spawning, taken by Kei Sasaki (Betta
House)
Picture 6: B. coccina spawning, taken by Kei Sasaki (Betta
House)
Picture 7: B. coccina bubblenest, taken by Kei Sasaki (Betta
House)
Picture 8: B. coccina male, taken by Arend van den Nieuwenhuizen
Picture 9: B. coccina male, taken by Jirapat Chunthapong
Pictures of chapter 'other species' by: See chapter.
Special
thanks to
E. Naus for helping me obtain Betta coccina.
The rightful owners of the pictures.
References
Vierke, J. 1979. Betta coccina nov. spec., ein neuer Kampffisch
von Sumatra.
Aquarium Aqua Terra 288-289.
The
International Betta Congress - Species Maintenance Program website
and species list.
Ralph
Tran of the International Betta Congress - Species Maintenance
Program for tips and facts.
E.
Naus' website, Betta coccina page.
©
Stefan vd Voort 2002